to say, to tell somebody / to call, to name, to give a name / to cry, to call, to make a noise (such as an animal)

What? What did you say?

whatchamacallit, X

past tense and perfective aspect of 言う (iu, yū) [godan]

nonetheless, although, be that as it may

in other words; what I (you, etc.) mean is

to startle; to surprise; to astonish

an organism; a life form

upper, over

the start / a high place; the top / one with higher rank