to say, to tell somebody / to call, to name, to give a name / to cry, to call, to make a noise (such as an animal)

What? What did you say?

whatchamacallit, X

past tense and perfective aspect of 言う (iu, yū) [godan]

nonetheless, although, be that as it may

in other words; what I (you, etc.) mean is

to startle; to surprise; to astonish

what

What?, Huh?

what / that thing (used in place of words you can't remember at the moment.)