The fourth species, the nothospecies Fragaria × ananassa, was originally established for the hybrids F. chiloensis × F. virginiana, which had originated under cultivation and are known as the large-fruited garden strawberries. When the hybrid populations in the coastal regions of northwestern North America were identified as introgressants of the same two octoploid species, they were combined with the cultivated garden strawberries into the nothospecies F. ×ananassa.