Aedeagus in lateral view with preatrium well developed, almost as long as shaft, shaft slender, tubular, straight, diverging from line of preatrium about 30°, with membranous flanges on dorsal side variable in shape, dorsoatrium absent but with pair of ligaments connecting to anal tube laterally, in ventral view aedeagus shaft and preatrium nearly broadened at atrium, gonopore apical (Figs 4C, 4F–4I, 5A and 5G–5I ).