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Detailed studies have shown that LmrP mediates the transbilayer movement of the amphiphilic cationic substrate TMA-DPH (1-(4-(trimethylamino)phenyl)-6-phenylhexa-1,3,5-triene) from the inner leaflet of the cytoplasmic membrane to the external side of the cell [5 ], thus effectively facilitating the transbilayer efflux of substrate across a permeability barrier at a rate that exceeds the rate of passive substrate influx.
In 2013, Tominaga and co–workers crystallized carbamoylated and cyanated forms of HypE from Thermococcus kodakarensis after in vitro incubation of the protein with potassium cyanide and ATP [27 ].
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